:root{--ink:#eee8dc;--muted:#aab2aa;--paper:#171d1a;--paper-2:#202722;--card:#1e2521;--line:#3f4942;--accent:#c8b890;--accent-2:#9f957b;--dark:#101512;--max:1180px;--shadow:0 12px 34px rgba(0,0,0,.28);--frame:rgba(245,245,240,.62)}
*{box-sizing:border-box}html{scroll-behavior:smooth}body{margin:0;background:radial-gradient(circle at 78% 3%,#242d28 0,#171d1a 35%,#111613 100%);color:var(--ink);font-family:Arial,Helvetica,sans-serif;line-height:1.62}a{color:inherit}img{max-width:100%;display:block}button,input{font:inherit}em{font-family:Georgia,"Times New Roman",serif}
.site-header{position:sticky;top:0;z-index:50;background:rgba(23,29,26,.96);backdrop-filter:blur(8px);border-bottom:1px solid var(--line)}.header-inner{max-width:var(--max);margin:auto;display:flex;align-items:center;justify-content:space-between;gap:26px;padding:14px 24px}.brand{text-decoration:none;display:flex;align-items:center;gap:14px;min-width:max-content}.brand-mark{width:38px;height:38px;border:1px solid var(--accent);border-radius:50%;display:grid;place-items:center;color:var(--accent);font:700 18px/1 Arial,sans-serif}.brand-text strong{display:block;font:500 18px/1.1 Georgia,serif;letter-spacing:.11em;text-transform:uppercase}.brand-text span{display:block;margin-top:3px;color:var(--muted);font:11px/1.2 Arial,sans-serif;letter-spacing:.08em;text-transform:uppercase}.nav{display:flex;align-items:center;gap:20px;font:13px/1 Arial,sans-serif}.nav a{text-decoration:none;padding:10px 0;border-bottom:2px solid transparent;white-space:nowrap}.nav a:hover,.nav a.active{border-color:var(--accent)}.menu-toggle{display:none;border:1px solid var(--line);background:var(--card);color:var(--ink);padding:8px 10px;border-radius:4px}
h1,h2,h3{font-family:Georgia,"Times New Roman",serif;font-weight:400;line-height:1.14;margin-top:0}h1{font-size:clamp(42px,6vw,78px);letter-spacing:-.035em;margin-bottom:20px}h2{font-size:clamp(30px,4vw,46px);letter-spacing:-.02em;margin-bottom:18px}h3{font-size:24px;margin-bottom:10px}.eyebrow{font:700 12px/1 Arial,sans-serif;letter-spacing:.16em;text-transform:uppercase;color:var(--accent);margin-bottom:18px}.lead{font:21px/1.55 Georgia,"Times New Roman",serif;color:#d7d4ca;max-width:760px}.muted{color:var(--muted)}
.hero{max-width:var(--max);margin:0 auto;padding:68px 24px 54px;display:grid;grid-template-columns:1.08fr .92fr;gap:50px;align-items:center}.hero-actions{display:flex;gap:12px;flex-wrap:wrap;margin-top:28px}.btn{display:inline-block;text-decoration:none;font:700 13px/1 Arial,sans-serif;letter-spacing:.04em;padding:13px 17px;border-radius:3px;border:1px solid var(--accent);background:var(--accent);color:#171d1a}.btn.secondary{background:transparent;color:var(--accent)}.stats{display:grid;grid-template-columns:repeat(3,1fr);gap:1px;background:var(--line);border:1px solid var(--line);margin-top:38px}.stat{background:var(--card);padding:22px}.stat strong{display:block;font:400 32px Georgia,serif;color:var(--accent)}.stat span{font:11px/1.4 Arial,sans-serif;color:var(--muted);text-transform:uppercase;letter-spacing:.07em}.hero-random-wrap{min-width:0}.hero-random-grid{display:grid;grid-template-columns:1fr 1fr;gap:13px}.random-photo{margin:0;min-width:0}.random-photo img{width:100%;height:245px;object-fit:contain;background:#0f1411;border:1.75px solid var(--frame);padding:5px 8px;box-shadow:var(--shadow)}.random-photo figcaption{font:12px/1.35 Arial,sans-serif;color:var(--muted);margin-top:6px;min-height:32px}.random-hint{font:11px/1.4 Arial,sans-serif;color:#89928b;margin:10px 0 0;text-align:right}
.section{padding:68px 24px;border-top:1px solid var(--line)}.section-inner{max-width:var(--max);margin:auto}.section-head{display:grid;grid-template-columns:.7fr 1.3fr;gap:48px;margin-bottom:34px}.section-head p{margin:0;color:#bec5be;font:18px/1.62 Georgia,"Times New Roman",serif}.card-grid{display:grid;grid-template-columns:repeat(3,1fr);gap:20px}.card{background:linear-gradient(145deg,var(--card),#171e1a);border:1px solid var(--line);padding:26px;text-decoration:none;transition:.18s ease}.card:hover{transform:translateY(-3px);box-shadow:var(--shadow);border-color:#68736b}.card small{font:11px/1 Arial,sans-serif;color:var(--accent);text-transform:uppercase;letter-spacing:.1em}.card h3{margin-top:12px}.card p{color:var(--muted);margin-bottom:0}.archive-note{margin-top:34px;border:1px solid var(--line);background:#151b18;padding:15px 18px;color:#aeb8b0;font-size:13px}
.page-hero{max-width:var(--max);margin:auto;padding:62px 24px 40px}.page-hero h1{font-size:clamp(40px,5vw,64px)}.breadcrumb{font:12px/1 Arial,sans-serif;color:var(--muted);margin-bottom:24px}.breadcrumb a{text-decoration:none}.content-wrap{max-width:var(--max);margin:auto;padding:0 24px 80px}.prose-wide{max-width:850px}.prose-wide p{font:18px/1.7 Georgia,"Times New Roman",serif;color:#cbd2cc}.filterbar{display:flex;gap:12px;margin:0 0 28px}.search{width:100%;padding:13px 14px;border:1px solid var(--line);background:var(--card);color:var(--ink);outline:none}.search::placeholder{color:#89928b}.search:focus{border-color:var(--accent)}
.genus-grid{display:grid;grid-template-columns:repeat(4,1fr);gap:14px;align-items:stretch}.genus{background:var(--card);border:1px solid var(--line);padding:20px;min-height:112px;display:flex;flex-direction:column;justify-content:space-between}.genus strong{display:block;font:400 22px Georgia,serif;font-style:italic}.genus-meta{margin-top:12px;display:flex;justify-content:space-between;gap:10px;align-items:end;font:12px/1.3 Arial,sans-serif;color:var(--muted)}.genus-link{font-weight:700;color:var(--accent);text-decoration:none}.genus-link:hover{text-decoration:underline}.genus-ready{border-color:#6d746d}.genus-status{color:#8d978f}
.gallery-intro{max-width:790px}.gallery-detail-link{margin-top:22px;font-size:15px}.gallery-detail-link a{text-decoration:none;border-bottom:1px solid var(--line)}.copyright-notice{border:1px solid var(--line);background:var(--card);padding:14px 16px;margin:0 0 30px;color:#cbd0cb;font-size:14px}.copyright-notice strong{color:var(--accent)}.species-gallery{display:grid;grid-template-columns:repeat(4,minmax(0,1fr));gap:30px 18px;margin-top:10px}.species-photo{margin:0;min-width:0}.species-photo button{display:block;width:100%;padding:0;border:0;background:none;cursor:zoom-in;text-align:inherit}.species-photo img{width:100%;aspect-ratio:4/3;object-fit:contain;background:var(--paper-2);border:1.75px solid var(--frame);padding:5px 8px;box-shadow:0 5px 18px rgba(0,0,0,.16);transition:transform .16s ease,box-shadow .16s ease}.species-photo button:hover img{transform:translateY(-2px);box-shadow:var(--shadow)}.species-photo figcaption{margin-top:8px;font:15px/1.3 Georgia,serif}.lightbox{position:fixed;inset:0;z-index:80;display:none;align-items:center;justify-content:center;padding:30px 82px 24px;background:rgba(8,11,9,.96)}.lightbox.open{display:flex}.lightbox-inner{max-width:min(1400px,92vw);max-height:95vh;display:flex;flex-direction:column;align-items:center}.lightbox img{max-width:100%;max-height:82vh;width:auto;height:auto;border:1.75px solid var(--frame);padding:6px 9px;background:#111613;box-shadow:0 18px 60px rgba(0,0,0,.42)}.lightbox-meta{display:flex;justify-content:center;gap:18px;margin-top:10px}.lightbox-count{font:12px Arial,sans-serif;color:#c8cec9}.lightbox-close{position:fixed;top:18px;right:22px;border:1px solid rgba(255,255,255,.48);background:rgba(0,0,0,.38);color:#fff;width:46px;height:46px;border-radius:50%;font:31px/1 Arial;cursor:pointer}.lightbox-prev,.lightbox-next{position:fixed;top:50%;transform:translateY(-50%);border:0;background:rgba(0,0,0,.28);color:#fff;width:54px;height:72px;font:48px/1 Arial;cursor:pointer}.lightbox-prev{left:16px}.lightbox-next{right:16px}
.topic-grid{display:grid;grid-template-columns:repeat(2,1fr);gap:20px}.topic-card{background:var(--card);border:1px solid var(--line);padding:26px}.topic-card h2{font-size:28px}.topic-card p{color:#bcc6be}.related{margin-top:52px}.simple-list{display:grid;grid-template-columns:repeat(2,1fr);gap:16px}.list-card{border-top:1px solid var(--line);padding:18px 0}.link-card{text-decoration:none}.link-card:hover h3{color:var(--accent)}.list-card h3{margin:0 0 8px;font-size:22px}.list-card p{margin:0;color:var(--muted)}.text-link{color:var(--accent);text-decoration:none;border-bottom:1px solid #5e665f}.imprint-card{max-width:800px;background:var(--card);border:1px solid var(--line);padding:30px}.imprint-card h2{font-size:26px;margin-top:28px}.imprint-card h2:first-child{margin-top:0}.notice{background:#202820;border-left:4px solid var(--accent);padding:17px 19px;margin-top:24px;color:#bcc5be}
/* Article overview */.article-shell{width:min(1180px,calc(100% - 42px));margin:auto}.article-shell>.hero{display:grid;grid-template-columns:1.5fr .7fr;gap:36px;padding:60px 0 36px}.article-shell>.hero h1{font-size:54px}.article-shell>.hero p{max-width:800px;color:#c4cec7;font:19px/1.6 Georgia,serif}.hero-note{border:1px solid var(--line);background:rgba(24,32,29,.7);padding:22px;align-self:end;color:var(--muted);font:14px/1.55 Arial,sans-serif}.hero-note strong{display:block;color:var(--text);margin-bottom:8px}.controls{display:flex;gap:10px;flex-wrap:wrap;align-items:center;border-top:1px solid var(--line);border-bottom:1px solid var(--line);padding:18px 0;margin-bottom:30px}.filter{border:1px solid var(--line);background:transparent;color:var(--muted);padding:9px 13px;border-radius:999px;cursor:pointer}.filter.active,.filter:hover{background:var(--paper-2);color:#fff}.controls .search{margin-left:auto;width:auto}.controls .search input{background:#0e1311;border:1px solid var(--line);color:#fff;padding:10px 14px;min-width:230px;border-radius:7px}.grid{display:grid;grid-template-columns:repeat(2,minmax(0,1fr));gap:22px;padding-bottom:70px}.grid .card{display:grid;grid-template-columns:245px 1fr;min-height:285px;padding:0;overflow:hidden}.pic{display:flex;align-items:center;justify-content:center;background:#090c0b;border-right:1px solid var(--line);padding:10px}.pic img{max-width:100%;max-height:265px;width:auto;height:auto;object-fit:contain;border:1.75px solid var(--frame);padding:4px 6px}.content{padding:24px;display:flex;flex-direction:column}.content h2{font-size:27px;line-height:1.12;margin:7px 0}.meta{font:700 12px Arial,sans-serif;text-transform:uppercase;letter-spacing:.12em;color:var(--accent)}.latin{font-style:italic;color:#d2dccd;margin-bottom:12px}.teaser{font:15px/1.55 Arial,sans-serif;color:#c3cdc6}.bottom{margin-top:auto;padding-top:16px;border-top:1px solid #2c3732;display:flex;gap:14px;align-items:end}.pub{font:12px/1.5 Arial,sans-serif;color:#89988f}.openbtn,.backbtn{margin-left:auto;display:inline-block;text-decoration:none;background:var(--accent);color:#111713;padding:10px 14px;border-radius:4px;font:700 13px Arial,sans-serif}.empty{display:none;color:var(--muted);padding:30px}
/* Full articles */.article-head{padding:58px 0 30px;border-bottom:1px solid var(--line)}.article-head h1{font-size:48px}.subtitle{font-style:italic;color:#d2dccd;font-size:20px}.article-meta{font:14px Arial,sans-serif;color:var(--muted);margin-top:12px}.article-wrap{display:grid;grid-template-columns:minmax(0,760px) 300px;gap:48px;padding:38px 0 72px}.fulltext{max-width:760px}.fulltext p{font:17px/1.72 Georgia,serif;color:#cbd4ce;margin:0 0 1.15em;hyphens:auto}.fulltext h2{font-size:25px;line-height:1.25;margin:2em 0 .65em;color:#edf2ed;border-top:1px solid #2b3731;padding-top:18px}.fulltext h2:first-child{margin-top:.15em;border-top:0}.fulltext .fig{margin:30px 0 36px;padding:14px 16px 12px;border:1.75px solid var(--frame);background:#0b0f0d;box-shadow:0 10px 25px rgba(0,0,0,.23)}.fulltext .fig img{max-width:100%;max-height:760px;width:auto;height:auto;margin:auto}.fulltext .fig figcaption{font:13px/1.5 Arial,sans-serif;color:#aab7ae;margin-top:11px}.sidebar{position:sticky;top:95px;align-self:start}.sidebox{border:1px solid var(--line);background:var(--card);padding:18px;margin-bottom:16px;font:14px/1.55 Arial,sans-serif;color:var(--muted)}.sidebox strong{display:block;color:var(--ink);margin-bottom:7px}.table-scroll{overflow-x:auto;margin:28px 0 36px;border:1px solid var(--line)}.article-table{width:100%;border-collapse:collapse;font:14px/1.45 Arial,sans-serif;color:#cbd4ce}.article-table th,.article-table td{padding:10px 12px;border-right:1px solid var(--line);border-bottom:1px solid var(--line);text-align:left}.article-table th{background:var(--card);color:#edf2ed}.article-back{margin-top:45px!important;padding-top:24px;border-top:1px solid var(--line)}.article-back .backbtn{margin-left:0}.article-lightbox{position:fixed;inset:0;background:rgba(0,0,0,.94);display:none;align-items:center;justify-content:center;z-index:120;padding:45px}.article-lightbox.open{display:flex}.article-lightbox img{max-width:94vw;max-height:88vh;border:1.75px solid var(--frame);padding:6px;background:#111}.article-lightbox-close{position:absolute;right:24px;top:18px;background:none;border:0;color:#fff;font-size:42px;cursor:pointer}.article-lightbox-cap{position:absolute;bottom:14px;color:#ddd;font:13px Arial,sans-serif}
.site-footer{background:var(--dark);color:#e9e7de;padding:40px 24px;border-top:1px solid var(--line)}.footer-inner{max-width:var(--max);margin:auto;display:grid;grid-template-columns:1fr auto;gap:30px;align-items:end}.footer-inner p{margin:0;color:#aab2aa;font:13px/1.6 Arial,sans-serif}.footer-links{display:flex;gap:18px;font:13px/1 Arial,sans-serif}.footer-links a{text-decoration:none}
@media(max-width:1080px){.nav{gap:14px;font-size:12px}.random-photo img{height:215px}.genus-grid{grid-template-columns:repeat(3,1fr)}}
@media(max-width:900px){.menu-toggle{display:block}.nav{display:none;position:absolute;left:0;right:0;top:67px;background:var(--paper);border-bottom:1px solid var(--line);padding:15px 24px;flex-direction:column;align-items:flex-start}.nav.open{display:flex}.hero{grid-template-columns:1fr;gap:34px;padding-top:48px}.hero-random-wrap{max-width:760px}.section-head{grid-template-columns:1fr;gap:8px}.card-grid,.topic-grid{grid-template-columns:1fr 1fr}.stats{grid-template-columns:1fr}.species-gallery{grid-template-columns:repeat(3,1fr)}.grid{grid-template-columns:1fr}.article-wrap{grid-template-columns:1fr}.sidebar{position:static}.article-shell>.hero{grid-template-columns:1fr}.footer-inner{grid-template-columns:1fr}}
@media(max-width:720px){.species-gallery{grid-template-columns:repeat(2,1fr)}.genus-grid{grid-template-columns:repeat(2,1fr)}.random-photo img{height:210px}.lightbox{padding:60px 50px 20px}.simple-list{grid-template-columns:1fr}}
@media(max-width:620px){.header-inner{padding:12px 16px}.brand-text span{display:none}.hero,.page-hero,.content-wrap,.section{padding-left:16px;padding-right:16px}.hero-random-grid{gap:9px}.random-photo img{height:180px;padding:4px 6px}.card-grid,.topic-grid{grid-template-columns:1fr}.grid .card{grid-template-columns:1fr}.pic{border-right:0;border-bottom:1px solid var(--line)}.article-shell{width:min(100% - 26px,1180px)}.article-shell>.hero{padding-top:42px}.article-shell>.hero h1{font-size:42px}.controls .search{margin-left:0;width:100%}.controls .search input{width:100%}.article-head h1{font-size:40px}.lead{font-size:18px}}
@media(max-width:430px){.hero-random-grid{grid-template-columns:1fr 1fr}.random-photo img{height:145px}.random-photo figcaption{font-size:11px}.species-gallery{grid-template-columns:1fr 1fr;gap:20px 10px}.genus-grid{grid-template-columns:1fr}.lightbox{padding:58px 12px 18px}.lightbox-prev,.lightbox-next{top:auto;bottom:18px;transform:none;width:46px;height:46px}.lightbox-prev{left:12px}.lightbox-next{right:12px}.lightbox-close{top:8px;right:8px}}

/* Unaufdringlicher globaler Seitenaufruf-Zähler */
.page-counter{color:var(--muted,#a9aaa6);font-size:.92em;white-space:nowrap}
.page-counter[hidden]{display:none!important}

/* Familien-Navigation der Bildsammlung */
.family-grid{display:grid;grid-template-columns:repeat(2,minmax(0,1fr));gap:22px;max-width:980px}
.family-card{display:block;text-decoration:none;background:linear-gradient(145deg,var(--card),#171e1a);border:1px solid var(--line);padding:30px;transition:.18s ease}
.family-card:hover{transform:translateY(-3px);box-shadow:var(--shadow);border-color:#68736b}
.family-card small{font:11px/1 Arial,sans-serif;color:var(--accent);text-transform:uppercase;letter-spacing:.1em}
.family-card h2{font-size:34px;margin:11px 0 12px}
.family-card p{color:var(--muted);margin:0 0 18px}
.family-card span{color:var(--accent);font-weight:700;font-size:13px}
.imprint-missing{border:1px solid #8d7356;background:#2b241d;padding:14px 16px;color:#ead8bf}
@media(min-width:901px){.nav{gap:15px;font-size:12px}}
@media(max-width:720px){.family-grid{grid-template-columns:1fr}}


/* Language selector and site-wide search */
.site-header{position:relative}
.header-inner{flex-wrap:nowrap}
.site-search-toggle{border:1px solid var(--line);background:transparent;color:var(--ink);font:12px/1 Arial,sans-serif;padding:8px 10px;cursor:pointer;white-space:nowrap}
.site-search-toggle:hover{border-color:var(--accent);color:#fff}
.language-bar{max-width:var(--max);margin:0 auto;padding:0 24px 8px;display:flex;justify-content:flex-end;gap:10px;font:12px/1.2 Arial,sans-serif}
.lang-link{opacity:.65;text-decoration:none;padding:3px 5px;border-bottom:1px solid transparent}
.lang-link:hover,.lang-link.active{opacity:1;border-bottom-color:var(--accent)}
.site-search-panel{position:fixed;inset:0;background:rgba(0,0,0,.76);z-index:180;padding:90px 18px 30px;overflow:auto}
.site-search-panel[hidden]{display:none!important}
.site-search-box{position:relative;max-width:820px;margin:0 auto;background:#111713;border:1px solid #647168;box-shadow:0 25px 80px rgba(0,0,0,.55);padding:26px}
.site-search-box label{display:block;font:600 14px/1.2 Arial,sans-serif;color:var(--ink);margin-bottom:10px}
.site-search-box input{width:100%;box-sizing:border-box;background:#080b09;border:1px solid #556059;color:#eef2ef;padding:13px 14px;font:16px/1.2 Arial,sans-serif;outline:none}
.site-search-box input:focus{border-color:var(--accent)}
.site-search-close{position:absolute;right:10px;top:6px;border:0;background:transparent;color:#dfe5df;font-size:30px;cursor:pointer}
.site-search-status{font:12px/1.5 Arial,sans-serif;color:var(--muted);margin:12px 0 4px}
.site-search-results{display:grid;gap:8px;margin-top:10px}
.search-result{display:block;text-decoration:none;border:1px solid var(--line);background:var(--card);padding:13px 15px}
.search-result:hover{border-color:#718077;background:#1a211d}
.search-result strong{display:block;color:#f1f4f1;font:600 15px/1.25 Arial,sans-serif;margin-bottom:4px}
.search-result span{display:block;color:var(--muted);font:12px/1.45 Arial,sans-serif}
@media(max-width:1080px){.site-search-toggle{font-size:0}.site-search-toggle::first-letter{font-size:16px}.language-bar{justify-content:flex-start;padding-left:24px}}
@media(max-width:900px){.site-search-toggle{margin-left:auto;margin-right:8px}.language-bar{padding:0 16px 8px}.site-search-panel{padding-top:70px}.site-search-box{padding:22px 16px}}